NEW YORK — At Bethel Gospel Assembly, a Pentecostal church in Harlem, Sundays once typically began with handshakes, hugs and kisses. The pastor welcomed parishioners from the pulpit to the 2,000-seat sanctuary — his greeting simultaneously relayed in French, Spanish and Italian by translators to the mostly European tourists crowded into the church’s balcony to hear the choirs sing.

In Harlem, a neighborhood with hundreds of churches — one of which, Elmendorf Reformed Church, dates to 1660 — choirs aren’t relegated to backing up the pastor. They are a primary attraction for church members and tourists alike. Many of the local gospel choirs are well known internationally — the topic of must-see travel lists — and before the pandemic they drew hundreds of visitors each week.

That changed the week of March 20, 2020, as pandemic closures went into effect. Church leaders including Bishop Carlton T. Brown, Bethel’s senior pastor for the past 21 years, told congregants that church doors were closing. Until further notice, services would be streamed online. Which also meant the choirs could no longer gather in person to sing.

“Some would say that in the United States, you can never shut the church down, the church will always be open here,” Brown said. “Then all of a sudden we came to a place where the church had to shut down.”

As services moved online, churches tried different approaches to including music. Some streamed soloists singing from home; others asked choir members to record themselves singing, then combined those videos to play during services.

At Bethel, the church invested in streaming services and saw an increase in attendance of Sunday and midweek services, with people searching for community during difficult times. The church’s online Bible studies have gone from 20 to 30 students each to well over 75 in each class. But many members spoke of missing the music, which is so central to the worship service.

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For the choir, singing online was technically challenging; the experience felt disjointed and the results were often disappointing. For churchgoers, watching services from home was lonely. They were grateful for the connection, but it wasn’t what it once was. They longed to pray and sing together and they missed the shared experience of worshipping — in particular, the experience of hearing their church choirs.

“I remember feeling so awkward not being in the house of God and not actually being able to sing on this pulpit,” said Warren S.L. Hicks, a choir worship leader who has been singing at Bethel since he was a boy. “It was almost like air being sucked out of my lungs.”

Leaders at these Black churches said that cutting music out of services was never an option.

You can’t have worship completely without music in the Black church,” said the Rev. Calvin Otis Butts III at the Abyssinian Baptist Church on West 138th Street. “It’s impossible. The Black church is culture bearer, so if you want to have the best of poetry, the best of music, the best of our cultural expressions, you come to the church.”

During the pandemic, being separated from other church members created a sense of loneliness for congregants and choir members. No number of Zoom and FaceTime calls, Facebook Bible studies and streamed Sunday services could make up for not being able to hug someone in person, read a passage from Scripture, hold hands and come together to sing and worship.

For many, church music had become a source of strength and a lifeline. So reestablishing that lifeline became a top church priority.

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Church leaders in majority Black and Latino Harlem were aware that their community was being affected disproportionately by COVID-19, so they were especially sensitive about the risks of bringing people together again. At Canaan Baptist Church of Christ on 116th Street, more than 40 members of the congregation, including some choir members, died from COVID in 2020.

But church leaders also found that during a time of fear and uncertainty, churchgoers felt cut off from a source of strength they needed: their community of faith. Not being able to gather and sing together — at funerals, in particular — made the distance from one another feel vast. So they looked for ways to bring live music back.

At the onset of the pandemic, Abyssinian Baptist Church, founded in 1808, began streaming virtual services and Ramón Bryant Braxton, the director of music, asked soloists to come sing alone with musicians in the empty sanctuary.

By late April 2020, Braxton had another idea: He assembled additional groups of singers, including a quartet and an ensemble of nine. The groups would sing while socially distanced in the empty church on alternating Sundays.

“At one point, we rehearsed in the sanctuary and lined the balcony with the big group of nine spread out,” Braxton said. “But the challenge was that, because everyone was so far apart, they had to work twice as hard to hear each other. To create the sense of ensemble you can’t be in a vacuum.”

B. Dexter Allgood, minister of music at Canaan Baptist, said that his church typically had seven different choirs consisting of dozens of singers and that on any given Sunday before the pandemic, as many as 100 people would sing.

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“You have the word and then you have the word in song — there’s a connection there, and when it’s not there it feels like a void,” Allgood said, adding that he doesn’t think that before the pandemic he had ever spent a single Sunday in his adult life away from a sanctuary.

After trying various methods of keeping music in the service virtually, Allgood pulled together a group of eight singers to come sing live from the sanctuary. Some of them stood socially distanced on the stage as others would sing while arranged among the empty pews.

Reuniting to sing together, even while distanced, allowed choir members to once again feel a sense of community.

“You think people get together because we all like to sing? Not necessarily,” said Frederick Myers, a singer at Canaan who also sings professionally. “They get together because they need somebody to talk to. It’s more counseling sometimes. You have an outlet, and once you have that person in the choir that you can talk to, you can sing because there’s somebody there who cares about what happened to you today.”

Myers has Stage 3 cancer; he said church music, and choral music in particular, had kept him going over the past year.

Hicks has been singing at Bethel for 30 years. His parents and his wife, Adrienne, are involved in Bethel’s music ministry. Adrienne Hicks’ father directed the youth ensemble in the 1970s and ’80s, and her mother directed the children’s choir. She and Hicks met at the church 33 years ago.

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These kinds of family connections aren’t rare in churches and among their choir members. Choirs at Black churches often include family members of multiple generations, and in many cases, choir members became so close that they considered one another family.

During the week, Warren Hicks is a manager at the emergency department at NYC Health + Hospitals in Harlem. The early days of the pandemic took a toll.

“When I couldn’t be in the sanctuary on a Sunday and I was in work pushing dead bodies,” Warren Hicks said, “it was important for me to have something in my ear to keep me going, and those were the songs, songs of the Lord.”

On the recent Easter Sunday, Renay Peters Joubert, a professional singer who has been at Abyssinian Baptist Church off and on for 15 years, returned to the church to sing an arrangement by her husband, Joseph Joubert, who accompanied her on piano. Services at Abyssinian hadn’t yet opened to congregants, marking a second Easter away from the church for many worshippers.

“It’s my whole being, the church music,” Renay Joubert said. “It just gives me that sense of calm, and that’s the only thing that really got me through this pandemic.”

At Bethel in the fall, Brown — feeling the strain of isolation in the community — decided to bring back in-person services with limited attendees. Safety was a priority. Only 50 parishioners could be in the church, which typically seats 2,000. The church did a soft opening in October and opened with a few members in mid-November for its 103rd anniversary.

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As restrictions in the city have lifted and more people have been vaccinated, the churches are welcoming limited numbers of parishioners back into the pews.

“You don’t know how it felt to just walk into that sanctuary again,” said Rita Williams, 59, who has been singing in three of Canaan Baptist Church’s choirs for seven years and who works as an emergency room technician at Mount Sinai Hospital. “I just had this peace and comfort.”

The experience isn’t yet what it once was — elbow bumps have replaced hugs and kisses; singers don’t share microphones; masks have to stay on. Most churches have changed how they observe communion. Those coming to church on Sunday have to register online and provide declarations ensuring that they are not displaying coronavirus symptoms. Once they arrive, temperatures are scanned and stickers displayed on pews to maintain social distance.

But it’s all worth it for a chance to worship together again.

“We need to share our experience and how we got through this,” Williams said. “We need that support from one another.”

The pews are not yet full, and the choirs aren’t back to full strength. But the churches are once again hosting gospel music and the churchgoers who missed it.

“I still miss looking out and seeing everyone singing in the pews, and I miss when the Holy Spirit come down,” Williams said. “I’m waiting for that day when the doors can open and we can all come in shouting and singing, ‘Hallelujah, praise God, thank you, Jesus’ — especially those of us who made it through the pandemic.”